2. Welling LLM & Shackelford TK (in press). The Oxford Handbook of Evolutionary Psychology and Behavioral Endocrinology. Oxford University Press: New York, NY.
1. Zeigler-Hill V, Welling LLM, & Shackelford TK (2015). Evolutionary Perspectives on Social Psychology. Springer Publishing: New York, NY. Purchase
Book Chapters
9. Mogilski JK, Wysocki A, Reeve SD, Mitchell V, Lunge J, & Welling LLM (in press). Stress hormones, physiology, and behavior. In LLM Welling & TK Shackelford (Eds.), The Oxford Handbook of Evolutionary Psychology and Behavioral Endocrinology. Oxford Publishing: Oxford, UK.
8. Welling LLM & Burriss RP (in press). Investigating the ovulatory cycle: An overview of research and methods. In LLM Welling & TK Shackelford (Eds.), The Oxford Handbook of Evolutionary Psychology and Behavioral Endocrinology. Oxford Publishing: Oxford, UK.
7. Welling LLM & Shackelford TK (in press). Integrating mechanisms and functions to understand behavior. In LLM Welling & TK Shackelford (Eds.), The Oxford Handbook of Evolutionary Psychology and Behavioral Endocrinology. Oxford Publishing: Oxford, UK.
6. Welling LLM & Shackelford TK (in press). Future directions in human behavioral endocrinology. In LLM Welling & TK Shackelford (Eds.), The Oxford Handbook of Evolutionary Psychology and Behavioral Endocrinology. Oxford Publishing: Oxford, UK.
5. Nicolas SCA & Welling LLM (2015). The Darwinian mystique? Synthesizing evolutionary psychology and feminism. In V Zeigler-Hill, LLM Welling, & TK Shackelford (Eds.), Evolutionary Perspectives on Social Psychology (pp. 203-212). Springer Publishing: New York, NY. PDF
4. Welling LLM, Zeigler-Hill V, & Shackelford TK (2015). Integrating evolutionary psychology and social psychology: Reflections and future directions. In V Zeigler-Hill,LLM Welling, & TK Shackelford (Eds.), Evolutionary Perspectives on Social Psychology (pp. 493-500). Springer Publishing: New York, NY. PDF
3. Zeigler-Hill V, Welling LLM, & Shackelford TK (2015). How can an understanding of evolutionary psychology contribute to social psychology? In V Zeigler-Hill, LLM Welling, & TK Shackelford (Eds.), Evolutionary Perspectives on Social Psychology (pp. 3-11). Springer Publishing: New York, NY. PDF
2. Welling LLM (2014). Female Orgasm. In VA Weekes-Shackelford & TK Shackelford (Eds.), Evolutionary Perspectives on Human Sexual Psychology and Behavior (pp. 223-242). Springer Publishing: New York, NY. PDF
1. Welling LLM & Puts DA (2014). Female adaptations to ovulation. In VA Weekes-Shackelford & TK Shackelford (Eds.), Evolutionary Perspectives on Human Sexual Psychology and Behavior (pp. 243-260). Springer Publishing: New York, NY. PDF
Journal Articles
67. Brindley S, McDonald MM, Welling LLM, & Zeigler-Hill V. (in press). An evolutionary perspective on intergroup dating bias. Comprehensive Results in Social Psychology.
66. Pham MN, Barbaro N, Holub AM, Holden CJ, Mogilski JK, Lopes GS, Nicolas SCA, Sela Y, Shackelford TK, Zeigler-Hill V, & Welling LLM (in press). Do men produce higher-quality ejaculates when primed with thoughts of partner infidelity? Evolutionary Psychology. PDF
65. Welling LLM, Allan K, Bestelmeyer PEG, Jones BC, & DeBruine LM (2017). Sexual dimorphism effects upon neurophysiological correlates of face processing in women. Adaptive Human Behavior and Physiology, 3(4), 337-350. PDF
64. Donaldson SH, Welling LLM, & Reeve SD (2017). The influence of hormone replacement therapy on mating psychology among post-menopausal women. Personality and Individual Differences, 115, 13-18. PDF
63. Hill AK, Cárdenas RA, Wheatley JR, Welling LLM, Burriss RP, Claes P, Apicella CL, McDaniel MA, Little AC, Shriver MD, & Puts DA (2017). Are there vocal cues to human developmental stability? Relationships between facial fluctuating asymmetry and voice attractiveness. Evolution and Human Behavior, 38(2), 249-258.PDF
62. Reeve SD, Kelly K, & Welling LLM (2017). The effect of mate value feedback on women's mating aspirations and mate preference. Personality and Individual Differences, 115, 77-82. PDF
61. Mogilski JK, & Welling LLM (2017). The relative importance of sexual dimorphism, fluctuating asymmetry, and color cues to health during evaluation of potential partners' facial photographs: A conjoint analysis study. Human Nature, 28, 53-75. PDF
60. Mogilski JK, & Welling LLM (2017). Staying friends with an ex: Sex and dark personality traits predict motivations for post-relationship friendship. Personality and Individual Differences, 115, 114-119. PDF
59. Nicolas SCA, & Welling LLM (2017). Women's acceptance of cosmetic surgery across the menstrual cycle. Personality and Individual Differences, 115, 99-102. PDF
58. Bird BM, Welling LLM, Ortiz TL, Moreau BJP, Hansen S, Emond M, Goldfarb B, Bonin PL, & Carré JM (2016). Effects of exogenous testosterone and mating context on men's preferences for female facial femininity. Hormones and Behavior, 85, 76-85. PDF
57. Puts DA, Hill AK, Bailey D, Walker R, Rendall D, Wheatley JR, Welling LLM, Dawood K, Cárdenas RA, Burriss RP, Jablonski N, Shriver M, Weiss D, Lameira A, Apicella C, Owren MJ, Barelli C, Glenn M, & Ramos-Fernandez G (2016). Sexual selection on male vocal fundamental frequency in humans and other anthropoids. Proceedings of the Royal Society of London - Series B, 283(1829), 20152830.PDF
56. Reeve SD, Kelly K, & Welling LLM (2016). Transitory environmental threat alters sexually dimorphic mate preferences and sexual strategy. Evolutionary Psychological Science, 2, 101-113. PDF
55. Welling LLM (2016). Synthetic hormone dose in hormonal contraceptives predicts individual differences in personality. Social Behavior Research and Practice, 1(1), 13-16. PDF
54. Welling LLM, Moreau BJP, Bird BM, Hansen S, & Carré JM (2016). Exogenous testosterone increases men's perceptions of their own physical dominance. Psychoneuroendocrinology, 64, 136-142. PDF
53. Gangestad SW, Haselton MG, Welling LLM, Gildersleeve K, Pillsworth EG, Burriss RP, Larson CM, & Puts DA (2016). How valid are assessments of conception probability in ovulatory cycle research? Evaluations, recommendations, and theoretical implications. Evolution and Human Behavior,37(2), 85-96. PDFSupplementary Material
52. Mogilski JK, Memering S, Welling LLM, & Shackelford TK (2017). Monogamy and consensual non-monogamy: Alternative approaches to pursuing a strategically pluralistic mating strategy. Archives of Sexual Behavior, 46, 407-417. PDF
51. McGuire MC, Williams KL, Welling LLM, & Vonk J (2015). Cognitive bias in rats is not influenced by oxytocin. Frontiers in Psychology, 6, 1306-1331. PDF
50. Gonzalez-Santoyo I, Wheatley JR, Welling LLM, Cardenas RA, Jimenez-Trejo F, Dawood K, & Puts DA (2015). The face of female dominance: Women with dominant faces have lower cortisol. Hormones and Behavior, 71, 16-21. PDF
49. McCoy MG, Welling LLM, & Shackelford TK (2015). Development and initial psychometric assessment of the Reasons for Pretending Orgasm Inventory. Evolutionary Psychology,13(1), 89-101. PDF
48. Puts DA, Pope LE, Hill AK, Cardenas RA, Welling LLM, Wheatley JR, & Breedlove SM (2015). Fulfilling desire: Evidence for negative feedback between men's testosterone, sociosexual psychology, and sexual partner number. Hormones & Behavior, 70, 14-21. PDF
47. Li Y, Bailey DH, Winegard B, Puts DA, Welling LLM, & Geary DC (2014). Women's preference for masculine traits is disrupted by images of male-on-female aggression. PLoS One, 9(10), e110497. PDF
46. Doll LM, Hill AK, Cárdenas RA, Welling LLM, Wheatley JR, & Puts DA (2014). How well do men’s faces and voices index mate quality and dominance? Human Nature, 25, 200-212. PDF
45. Mogilski J, Wade TJ, & Welling LLM (2014). Prioritization of potential mates' history of sexual fidelity during a conjoint ranking task. Personality and Social Psychology Bulletin, 40(7), 884-889. PDF
44. Wheatley JR, Apicella CL, Burriss RP, Cardenas RA, Bailey D, Welling LLM, & Puts DA (2014). Women's faces and voices are cues to reproductive potential in industrial and forager societies. Evolution and Human Behavior, 35, 264-271. PDF
43. Hill AK, Hunt J, Welling LLM, Cardenas RA, Rotella MA, Wheatley JR, Dawood K, Shriver MD, & Puts DA (2013). Quantifying the strength and form of sexual selection on men’s traits. Evolution and Human Behavior, 34(5), 334-341. PDF
42. Jones BC, Fincher CL, Welling LLM, Little AC, Feinberg DR, Watkins CD, Al-Dujaili EAS, & DeBruine LM (2013). Salivary cortisol and pathogen disgust predict men's preferences for feminine shape cues in women's faces. Biological Psychology, 92, 233-240. PDF
41. Pham MN, Shackelford TK, Sela Y, & Welling LLM (2013). Is cunnilingus-assisted orgasm a male sperm-retention strategy? Evolutionary Psychology,11(2), 405-414. PDF
40. Pham MN, Shackelford TK, Welling LLM, Ehrke AD, Sela Y, & Goetz AT (2013). Oral sex, semen displacement, and sexual arousal: Testing the ejaculate adjustment hypothesis. Evolutionary Psychology,11(5), 1130-1139. PDF
39. Puts DA, Bailey DH, Cárdenas RA, Burriss RP, Welling LLM, Wheatley JR, & Dawood K (2013). Women's attractiveness changes with estradiol and progesterone across the ovulatory cycle. Hormones and Behavior, 63, 13-19. PDF
38. Tinlin RM, Watkins CD, Welling LLM, DeBruine LM, Al-Dujaili EA, & Jones BC (2013). Perceived facial adiposity conveys information about women’s health. British Journal of Psychology, 104, 235-248. PDF
37. Welling LLM (2013). Psychobehavioral effects of hormonal contraceptive use. Evolutionary Psychology, 11(3), 718-742. PDF
36. Welling LLM, Persola L, Wheatley JR, Cárdenas RA, & Puts DA (2013). Competition and men’s face preferences. Personality and Individual Differences, 54, 414-419. PDF
35. Welling LLM, Singh K, Puts DA, Jones BC, & Burriss RP (2013). Self-reported sexual desire in homosexual men and women predicts preferences for sexually dimorphic facial cues. Archives of Sexual Behavior, 42(5),785-791. PDF
34. Puts DA, Dawood K, & Welling LLM (2012). Why women have orgasms: An evolutionary analysis. Archives of Sexual Behavior, 41, 1127-1143. PDF
33. Puts DA, Welling LLM, Burriss RP, & Dawood K (2012). Men's masculinity and attractiveness predict their female partners' reported orgasm frequency and timing. Evolution and Human Behavior, 33(1), 1-9. PDF
32. Welling LLM, Puts DA, Roberts SC, Little AC, & Burriss RP (2012). Hormonal contraceptive use and mate retention behavior in women and their male partners. Hormones and Behavior, 61, 114-120. PDF
31. Burriss RP, Roberts SC, Welling LLM, Puts DA, & Little AC (2011). Heterosexual romantic couples mate assortatively for facial symmetry, but not masculinity. Personality and Social Psychology Bulletin, 37, 601-613. PDF
30. Burriss RP, Welling LLM, & Puts DA (2011). Mate-preference drives mate-choice: Men's self-rated masculinity predicts their female partner's preference for masculinity. Personality and Individual Differences, 51, 1023-1027. PDF
29. Burriss RP, Welling LLM, & Puts DA (2011). Men’s attractiveness predicts their preference for female facial femininity when judging for short-term, but not long-term, partners. Personality and Individual Differences, 50, 542-546. PDF
28. DeBruine LM, Jones BC, Little AC, Crawford JR, & Welling LLM (2011). Further evidence for regional variation in women's masculinity preferences. Proceedings of the Royal Society of London B, 278, 813-814. PDF
27. Jones BC, Little AC, Watkins CD, Welling LLM, & DeBruine LM (2011). Reported sexual desire predicts men’s preferences for sexually dimorphic cues in women’s faces. Archives of Sexual Behavior, 40(6), 1281-1285. PDF
26. Puts DA, Barndt JL, Welling LLM, Dawood K, & Burriss RP (2011). Intrasexual competition among women: Vocal femininity affects perceptions of attractiveness and flirtatiousness. Personality and Individual Differences, 50(1), 111-115. PDF
25. Vukovic J, Jones BC, Feinberg DR, DeBruine LM, Smith FG, Welling LLM, & Little AC (2011). Variation in perceptions of physical dominance and trustworthiness predicts individual differences in the effect of relationship context on women's preferences for masculine pitch in men's voices. British Journal of Psychology, 102, 37-48. PDF
24. Welling LLM, Burriss RP, & Puts DA (2011). Mate retention behavior modulates men’s preferences for self-resemblance in infant faces. Evolution and Human Behavior, 32(2), 118-126. PDF
23. Bestelmeyer PEG, Jones BC, DeBruine LM, Little AC, Schneider A, Welling LLM, & Conway CA (2010). Face aftereffects suggest interdependent processing of expression and sex and expression and race. Visual Cognition, 18, 255-274. PDF
22. DeBruine LM, Jones BC, Crawford JR, Welling LLM, & Little AC (2010). The health of a nation predicts their mate preferences: Cross-cultural variation in women's preferences for masculinized male faces. Proceedings of the Royal Society of London B, 277(1692), 2405-2410. PDF
21. DeBruine LM, Welling LLM, Jones BC, & Little AC (2010). Opposite effects of visual versus imagined presentation of faces on subsequent sex perception. Visual Cognition, 18, 816-828. PDF
20. Jones BC, DeBruine LM, Main JC, Little AC, Welling LLM, Feinberg DR, & Tiddeman BP (2010). Facial cues of dominance modulate the short-term gaze-cuing effect in human observers. Proceedings of the Royal Society of London B, 277, 617-624. PDF
19. Jones BC, Main JC, DeBruine LM, Little AC, & Welling LLM (2010). Reading the look of love: Sexually dimorphic cues in opposite-sex faces influence gaze-categorization. Psychological Science, 21, 796-798. PDF
18. Vukovic J, Jones BC, DeBruine LM, Feinberg DR, Smith FG, Little AC, Welling LLM, & Main JC (2010). Women’s own voice pitch predicts their preferences for masculinity in men’s voices. Behavioral Ecology, 21, 767-772. PDF
17. Smith FG, DeBruine LM, Jones BC, Krupp DB, Welling LLM, & Conway CA (2009). Attractiveness qualifies the effect of observation on trusting behavior in an economic game. Evolution and Human Behavior, 30, 393-397. PDF
16. Smith FG, Jones BC, Little AC, DeBruine LM, Welling LLM, Vukovic J, & Conway CA (2009). Hormonal contraceptive use and perceptions of trust modulate the effect of relationship context on women's preferences for sexual dimorphism in male face shape. Journal of Evolutionary Psychology, 7, 195-210. PDF
15. Smith FG, Jones BC, Welling LLM, Little AC, Vukovic J, Main JC, & DeBruine LM (2009). Waist-hip ratio predicts women's preferences for masculine male faces, but not perceptions of men's trustworthiness. Personality and Individual Differences, 47, 476-480. PDF
14. Vukovic J, Jones BC, DeBruine LM, Little AC, Feinberg DR, & Welling LLM (2009). Circum-menopausal effects on women's judgements of facial attractiveness. Biology Letters, 5, 62-64. PDF
13. Welling LLM, Bestelmeyer PEG, DeBruine LM, Little AC, Conway CA, & Jones BC (2009). View-contingent aftereffects suggest joint coding of face shape and view. Perception, 38, 133-141. PDF
12. Welling LLM, Jones BC, & DeBruine LM (2009). Extraversion predicts individual differences in women’s face preferences. Personality and Individual Differences, 47, 996-998. PDF
11. Bestelmeyer PEG, Jones BC, DeBruine LM, Little AC, Perrett DI, Schneider A, Welling LLM, & Conway CA (2008). Sex-contingent face aftereffects depend on perceptual category rather than structural encoding. Cognition, 107, 353-365. PDF
10. Conway CA, Jones BC, DeBruine LM, Little AC, Hay J, Welling LLM, Perrett DI & Feinberg DR (2008). Integrating physical and social cues when forming face preferences: Differences among low and high anxiety individuals. Social Neuroscience, 3, 89-95. PDF
9. Vukovic, J, Feinberg DR, Jones BC, DeBruine LM, Welling LLM, Little AC, & Smith FG (2008). Self-rated attractiveness predicts individual differences in women's preferences for masculine men's voices. Personality and Individual Differences, 45, 451-456. PDF
8. Welling LLM, Jones BC, & DeBruine LM (2008). Sex drive is positively associated with women's preferences for sexual dimorphism in men's and women's faces. Personality and Individual Differences, 44, 161-170. PDF
7. Welling LLM, Jones BC, DeBruine LM, Little AC, & Smith FG (2008). Exposure to sexually attractive men decreases women’s preferences for feminine faces. Journal of Evolutionary Psychology, 6, 219-230. PDF
6. Welling LLM, Jones BC, DeBruine LM, Smith FG, Feinberg DR, Little AC, & Al-Dujaili EAS (2008). Men report stronger attraction to femininity in women’s faces when their testosterone levels are high. Hormones and Behavior, 54, 703-708. PDF
5. Conway CA, Jones BC, DeBruine LM, Welling LLM, Law Smith MJ, Perrett DI, Sharpe M, & Al-Dujalli EA (2007). Salience of emotional displays of danger and contagion in faces is enhanced when progesterone levels are raised. Hormones and Behavior, 51, 202-206. PDF
4. Jones BC, DeBruine LM, Little AC, Conway CA, Welling LLM, & Smith FG (2007). Sensation seeking and men's face preferences. Evolution and Human Behavior, 28, 439-446. PDF
3. Welling LLM, Jones BC, DeBruine LM, Conway CA, Law Smith MJ, Little AC, Feinberg DR, Sharp MA, & Al-Dujaili EA (2007). Raised salivary testosterone in women is associated with increased attraction to masculine faces. Hormones and Behavior, 52, 156-161. PDF
2. Welling LLM, Conway CA, DeBruine LM, & Jones BC (2007). Perceived vulnerability to disease is positively related to the strength of preferences for apparent health in faces. Journal of Cultural and Evolutionary Psychology, 5, 131-139. PDF
1. Buckingham G, DeBruine LM, Little AC, Welling LLM, Conway CA, Tiddeman BP, & Jones BC (2006). Visual adaptation to masculine and feminine faces influences generalized preferences and perceptions of trustworthiness. Evolution and Human Behavior, 27, 381-389. PDF
Other Publications
26. Mogilski JK, & Welling LLM (2016). Honest signaling: A Review of Tucker Max & Geoffrey Miller, Mate. Evolutionary Psychological Science, 2(4), 374.PDF
25. Donaldson SH (in press). Mark Bekoff. In TK Shackelford & V Weekes-Shackelford (Eds.), The Encyclopedia of Evolutionary Psychological Science. Springer Publishing: New York, NY.
24. Donaldson SH (in press). Sarah Brosnan. In TK Shackelford & V Weekes-Shackelford (Eds.), The Encyclopedia of Evolutionary Psychological Science. Springer Publishing: New York, NY.
23. Lunge J (in press). Morality is Cooperation. In TK Shackelford & V Weekes-Shackelford (Eds.), The Encyclopedia of Evolutionary Psychological Science. Springer Publishing: New York, NY.
22. Lunge J (in press). Developmental Evidence. In TK Shackelford & V Weekes-Shackelford (Eds.), The Encyclopedia of Evolutionary Psychological Science. Springer Publishing: New York, NY.
21. Mitchell VE (in press). Reproduction. In TK Shackelford & V Weekes-Shackelford (Eds.), The Encyclopedia of Evolutionary Psychological Science. Springer Publishing: New York, NY.
20. Mitchell VE (in press). Medically-Assisted Reproduction. In TK Shackelford & V Weekes-Shackelford (Eds.), The Encyclopedia of Evolutionary Psychological Science. Springer Publishing: New York, NY.
19. Mogilski JK (in press). Science and Morality. In TK Shackelford & V Weekes-Shackelford (Eds.), The Encyclopedia of Evolutionary Psychological Science. Springer Publishing: New York, NY.
18. Mogilski JK (in press). Social Darwinism. In TK Shackelford & V Weekes-Shackelford (Eds.), The Encyclopedia of Evolutionary Psychological Science. Springer Publishing: New York, NY.
17. Nicolas SCA (in press). Genital mutilation. In TK Shackelford & V Weekes-Shackelford (Eds.), The Encyclopedia of Evolutionary Psychological Science. Springer Publishing: New York, NY.
16. Nicolas SCA (in press). Harris' moral landscape. In TK Shackelford & V Weekes-Shackelford (Eds.), The Encyclopedia of Evolutionary Psychological Science. Springer Publishing: New York, NY.
15. Nicolas SCA (in press). Sexism. In TK Shackelford & V Weekes-Shackelford (Eds.), The Encyclopedia of Evolutionary Psychological Science. Springer Publishing: New York, NY. 14. Reeve SD (in press). Better never to have been. In TK Shackelford & V Weekes-Shackelford (Eds.), The Encyclopedia of Evolutionary Psychological Science. Springer Publishing: New York, NY.
13. Reeve SD (in press). Game theory. In TK Shackelford & V Weekes-Shackelford (Eds.), The Encyclopedia of Evolutionary Psychological Science. Springer Publishing: New York, NY.
12. Reeve SD (in press). Measuring well-being. In TK Shackelford & V Weekes-Shackelford (Eds.), The Encyclopedia of Evolutionary Psychological Science. Springer Publishing: New York, NY.
11. Reeve SD (in press). Prisoner's dilemma. In TK Shackelford & V Weekes-Shackelford (Eds.), The Encyclopedia of Evolutionary Psychological Science. Springer Publishing: New York, NY.
10. Reeve SD (in press). Responsibility for child suffering. In TK Shackelford & V Weekes-Shackelford (Eds.), The Encyclopedia of Evolutionary Psychological Science. Springer Publishing: New York, NY.
9. Reeve SD (in press). The evolution of cooperation. In TK Shackelford & V Weekes-Shackelford (Eds.), The Encyclopedia of Evolutionary Psychological Science. Springer Publishing: New York, NY.
8. Welling LLM (in press). Hormone assays. In V Zeigler-Hill & TK Shackelford (Eds.)., The Encyclopedia of Personality and Individual Differences. Springer Publishing: New York, NY.
7. Welling LLM (in press). Judith Jarvis Thomson. In TK Shackelford & V Weekes-Shackelford (Eds.),The Encyclopedia of Evolutionary Psychological Science. Springer Publishing: New York, NY.
6. Welling LLM (in press). Ovulation. In TK Shackelford & V Weekes-Shackelford (Eds.),The Encyclopedia of Evolutionary Psychological Science. Springer Publishing: New York, NY.
5. Welling LLM (in press). Philippa Foot. In TK Shackelford & V Weekes-Shackelford (Eds.), The Encyclopedia of Evolutionary Psychological Science. Springer Publishing: New York, NY.
4. Wysocki A (in press). Debating procreation (with David Benatar). In TK Shackelford & V Weekes-Shackelford (Eds.), The Encyclopedia of Evolutionary Psychological Science. Springer Publishing: New York, NY.
3. Wysocki A (in press). Permissability of reproduction. In TK Shackelford & V Weekes-Shackelford (Eds.), The Encyclopedia of Evolutionary Psychological Science. Springer Publishing: New York, NY.
2. Wysocki A (in press). Would you kill the fat man? In TK Shackelford & V Weekes-Shackelford (Eds.), The Encyclopedia of Evolutionary Psychological Science. Springer Publishing: New York, NY.
1. Welling LLM (Dec. 2007). Time for T and masculine men? Testosterone levels linked to women’s preferences for masculine men. American Sexuality Magazine. National Sexuality Resource Center: San Francisco, CA.
